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Vertebrate
*** Shopping-Tip: Vertebrate
{{Taxobox begin | color = pink | name = Vertebrates}}
{{Taxobox image | image =
Image:Bluefin-big.jpg 250px|Northern Bluefin Tuna | caption =
Northern Bluefin Tuna, ''Thunnus thynnus''}}
{{Taxobox begin placement | color = pink}}
{{Taxobox regnum entry | taxon =
Animal Animalia}}
{{Taxobox phylum entry | taxon =
Chordata}}
{{Taxobox norank entry | taxon =
Craniata}}
{{Taxobox subphylum_entry | taxon = '''Vertebrata'''}}
{{Taxobox authority | author =
Georges Cuvier Cuvier | date = 1812}}
{{Taxobox end placement}}
{{Taxobox end}}
'''Vertebrata''' is a
subphylum of
chordates, specifically, those with backbones or
spinal columns. About 57,739
species of vertebrates have been described. [http://www.redlist.org/info/tables/table1]
Vertebrates started to evolve about 530 million years ago during the
Cambrian explosion, which is part of the
Cambrian period (first known vertebrate is
Myllokunmingia). The bones of the spinal column (or '''vertebral column''') are called
vertebrae. ''Vertebrata'' is the largest subphylum of chordates, and contains most animals with which people are generally familiar (except
insects).
Fish (including
lampreys, but traditionally not
hagfish, though this is now disputed),
Amphibia amphibians,
reptiles,
birds, and
mammals (including
humans) are vertebrates. Additional characteristics of the subphylum are a
muscular system that mostly consists of paired masses, as well as a
central nervous system which is partly located inside the backbone. Defining characteristics of a vertebrate are a
backbone or
spinal cord, a brain case, and an internal skeleton.
The internal
skeleton which defines vertebrates consists of
cartilage or
bone, or in some cases both. An outer skeleton in form of a bony armour was the first bony substance the vertebrates evolved. It is possible its primary function was as a phosphate reservoir, excreted as calcium phosphate and stored around the body, offering protection at the same time. The skeleton provides support to the organism during the period of growth. For this reason vertebrates can achieve larger sizes than
invertebrates, and on average vertebrates are in fact larger. The skeleton of most vertebrates, that is excluding the most primitive ones, consists of a
skull, the vertebral column and two pairs of
limbs. In some forms of vertebrates, one or both of these pairs of limbs may be absent, such as in
snakes or
whales. These limbs have been lost in the course of evolution.
The skull is thought to have facilitated the development of
Intelligence (trait) intelligence as it protects vital organs such as the brain, the eyes and the ears. The protection of these organs is also thought to have positively influenced the development of high responsiveness to the environment often found in vertebrates.
Both the vertebral column and the limbs support the body of the vertebrate overall. This support facilitates movement. Movement is normally achieved with muscles that are attached directly to the bones or cartilages. The contour of the body of a vertebrate is formed by the muscles. A skin covers the inner parts of a vertebrate's body. The
skin sometimes acts as a structure for protective features, such as horny scales or fur. Feathers may also be attached to the skin.
The trunk of a vertebrate is hollow and houses the internal organs. The heart and the respiratory organs are protected in the trunk. The heart is located behind the gills, or where there are lungs, in between the lungs.
The central nervous system of a vertebrate consists of the brain and the spinal cord. Both of these are characterized by being hollow. In lower vertebrates the brain mostly controls the functioning of the sense organs. In higher vertebrates the size of the brain relative to the size of the body is greater. This larger brain enables more intensive exchange of information between the different parts of the brain. The nerves from the spinal cord, which lies behind the brain, extend to the skin, the inner organs and the muscles. Some nerves are directly connected to the brain, linking the brain with the ears and lungs.
Vertebrates have been traced back to the
ostracoderms of the
Silurian Silurian Period (444 million to 409 million years ago) and the
conodonts, a group of eel-like vertebrates characterized by multiple pairs of bony toothplates.
Taxonomy
Classification after Benton, 2004 [http://palaeo.gly.bris.ac.uk/benton/vertclass.html#fishes].
* '''SUBPHYLUM VERTEBRATA'''
** Class '
Agnatha' (
paraphyletic)
*** Subclass â€
Conodonta
*** Subclass
Petromyzontidae (
lampreys)
*** Subclass â€
Pteraspidomorphi
*** Order â€
Thelodonti
*** Order â€
Anaspida
*** Subclass unnamed
**** Order â€
Galeaspida
**** Order â€
Pituriaspida
**** Order â€
Osteostraci
** '''Infraphylum
Gnathostomata''' (jawed vertebrates)
*** Class â€
Placodermi
*** Class
Chondrichthyes (cartilaginous fish)
*** Class â€
Acanthodii
*** Class
Osteichthyes (bony fish)
**** Subclass
Actinopterygii (ray-finned fish)
**** Subclass
Sarcopterygii (lobe-finned fish)
***** Order
Actinistia (
coelacanths)
***** Superclass
Tetrapoda (four-limbed vertebrates)
****** Class
amphibian Amphibia (amphibians)
****** Series
Amniota (amniotic embryo)
******* Class
Sauropsida (reptiles and birds)
******** Class
bird Aves (birds)
******* Class
Synapsida (mammal-like reptiles)
******** Class
mammal Mammalia (mammals)
See also
*
Invertebrate
External links
-
Tree of Life
-
Vertebrate Zoology
-
Vertebrate: individual or colony
{{Wikispecies|Vertebrata}}
Category:Chordates
ca:Vertebrat
cs:Obratlovci
cy:Fertebrat
da:Hvirveldyr
de:Wirbeltiere
es:Vertebrado
eo:Vertebrulo
fa:مهره‌داران
fr:Vertebrata
fy:Wringedier
ko:척추�물
id:Vertebrata
ia:Vertebrato
is:Hryggdýr
it:Vertebrata
he:בעלי חוליות
la:Vertebrata
lt:Stuburiniai
li:Gewèrvelde diere
ms:Vertebrat
nl:Gewervelden
ja:脊椎動物
nn:virveldyr
no:Virveldyr
oc:Categoria:Vertebrata
pl:Kręgowce
pt:Vertebrata
ru:Позвоночные
simple:Vertebrate
sk:Stavovce
sl:VretenÄ?arji
sr:Кичмењаци
fi:Selkärankaiset
sv:Ryggradsdjur
th:สัตว์มี�ระดู�สันหลัง
tr:Omurgalılar
zh:脊椎动物
*** Shopping-Tip: Vertebrate